Landlocked, lacustrine, pelagic, in upper water layers during summer and descending to deeper layers in winter. It spawns at 1-2 years, near shores above gravel or coarse sand, with most individuals spawning more than one season. Spawning starts when temperature reach about 19-20°C, in July to August . Adults form schools in the epilimnion during summer and tend to congregate in shallow waters near shores. They feed on plankton (cladocerans and copepods) and small fishes . It is threatened by water abstraction and pollution .
Dorsal spines (total): 3 - 4; Dorsal soft rays (total): 12 - 14; Anal spines: 3 - 4; Anal soft rays: 16 - 21. This species is distinguished from its congeners entering the freshwater in Mediterranean basin by the following characters: gill rakers 106-128; teeth on palatine and vomer well-developed, especially in juveniles .: fusiform / normal.
Behavioural data for Macedonia shad is currently being reviewed.
Deposits eggs near the shore where the sediment consists of gravel or coarse sand . Most individuals spawn in more than one season .
Freshwater; pelagic. Subtropical; 41°N - 40°N, 23°E - 24°E
plankton (cladocerans and copepods) and small fishes (Ref
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